The year is 1939 - it was then that the Neuroi appeared. Nobody knows where they came from or what their ultimate agenda is, but the fact remains that their attacks drove people out of their towns and cities. In order to take arms against them, humanity develops a new anti-Neuroi weapon called the "Striker Unit." Using the power of magic to fight against the monsters, this new device enhances and amplifies the power of female magic-wielders. To use this device, young witches from all over the world have been brought together to form an elite task force unit called the 501st Joint Fighter Wing, commonly known as the "Strike Witches."
福圓美里
as Yoshika Miyafuji
原由実
as Aleksandra Ivanovna Pokryshkin
門脇舞以
as Sanya V. Litvyak
小清水亜美
as Charlotte E. Yeager
川澄綾子
as Heinrike Prinzessin zu Sayn-Wittgenstein (voice)
水橋かおり
as Martina Crespi
森永理科
as Fernandia Malvezzi
名塚佳織
as Lynette Bishop
野川さくら
as Erica Hartmann
野水伊織
as Rosalie de Hemricourt de Grunne
大橋歩夕
as Eila Ilmatar Juutilainen
斎藤千和
as Francesca Lucchini