Fievel and his friend Tony Toponi find a map that they believe points to a treasure buried somewhere beneath Old New York, and the plucky rodent is determined to find it. However, what he discovers under the city is a tribe of Native American mice who were driven underground by prejudiced European immigrants.
Thomas Dekker
as Fievel Mousekewitz (voice)
Lacey Chabert
as Tanya Mousekewitz (voice)
Nehemiah Persoff
as Papa Mousekewitz (voice)
Erica Yohn
as Mama Mousekewitz (voice)
Dom DeLuise
as Tiger (voice)
Elaine Bilstad
as Cholena (voice)
Pat Musick
as Tony Toponi (voice)
René Auberjonois
as Dithering (voice)
David Carradine
as Chief Wulisso (voice)
Sherman Howard
as Police Chief McBrusque (voice)
Tony Jay
as Toplofty (voice)
Richard Karron
as O'Bloat (voice)