The story follows Ichika Hoshino, who hears a Miku song that she has never heard before at a CD shop and sees a Hatsune Miku whom she had never seen before. Ichika calls out to Miku, who surprised by the voice, disappears shortly after making eye contact. Miku later appears on Ichika's smartphone to tell her that she would like to reach others with her songs, but no matter how much she sings, her songs cannot reach them. After seeing Ichika reach others' hearts with her live street performance, Miku asks for her help.
藤田咲
as Miku Hatsune (voice)
下田麻美
as Rin Kagamine / Len Kagamine (voice)
浅川悠
as Luka Megurine (voice)
拝郷メイコ
as MEIKO (voice)
風雅なおと
as Kaito (voice)
野口瑠璃子
as Ichika Hoshino (voice)
礒部花凜
as Saki Tenma (voice)
上田麗奈
as Honami Mochizuki (voice)
中島由貴
as Shiho Hinomori (voice)
小倉唯
as Minori Hanasato (voice)
吉岡茉祐
as Haruka Kiritani (voice)
降幡愛
as Airi Momoi (voice)